Short answers, pulled from the story.
A sculptor named Nikolai Peristov found the left femur bone on the bank of the Irtysh River near Ust-Ishimsky District. He initially identified the material as mammoth ivory before noticing human characteristics upon closer inspection.
Carbon dating established that the fossil dates back 45,000 years ago. It stands as the oldest human fossil to receive such precise dating at the time of discovery.
The Y-DNA haplogroup K-M2308 defines the paternal lineage while his mitochondrial DNA belongs to haplogroup R*. Studies by Poznik et al. in 2016 identified him as K2(xK2b,K2c,K2d,K2e) with specific SNPs like M2308.
Ust'-Ishim man carries Neanderthal DNA in clusters instead of scattered pieces found in most modern non-African genomes. Genomic sequencing refined the estimated mating date between species to 52,000 and 58,000 years ago.
A 2016 study identified modern Tibetans as having the most alleles in common with Ust'-Ishim man. Another 2017 study found Siberian and East Asian populations shared 38% ancestry with him.