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— CH. 1 · INTRODUCTION —

Shrubland

6 min listen · Ch. 1 of 5
5 sections
  • Shrubland is one of the earth's most widespread and misunderstood landscapes, known by at least five different names depending on where you are standing: shrubland, scrubland, scrub, brush, or bush. The word itself only entered the language in 1903, yet the plant communities it describes have shaped continents for far longer. What makes a shrubland? Why do some of these places persist for centuries while others vanish after a single season? And why are they, despite their quiet appearance, some of the most fire-adapted environments on the planet? Those are the questions this documentary will explore.

  • Botanists draw the line at 8 metres. A woody plant with many stems that stays below that height is, by formal definition, a shrub rather than a tree. Within that broad category, ecologists in Australia developed a classification system based on structure, plant life-form, height, and foliage cover that has been widely adopted for mapping vegetation. Tall shrubs run from 2 to 8 metres. Small shrubs occupy the 1-to-2-metre band, and subshrubs stay below a single metre. Foliage cover matters just as much as height. A stand where canopy coverage runs from 70 to 100 percent is called closed-shrubland. Drop that coverage to between 30 and 70 percent and the same vegetation becomes open-shrubland. Below 30 percent the classification shifts again, down to tall shrubland and, at the sparsest end, tall open shrubland. For shorter growth under 2 metres, the same logic applies, yielding categories from closed-heath at the densest end to low open shrubland at the most sparse. In North America those same density bands carry the labels closed low shrubland and mid-dense low shrubland. The Australian system gave ecologists a shared vocabulary for comparing wildly different landscapes across the globe.

  • Chaparral in California, matorral in Chile, Mexico, and Spain, maquis in France and elsewhere around the Mediterranean, macchia in Italy, fynbos in South Africa, kwongan in Southwest Australia, caatinga in northeastern Brazil, cedar scrub in Texas Hill Country, Eastern Suburbs Banksia Scrub in Sydney. Each name points to the same basic type: dense thickets of evergreen sclerophyll shrubs and small trees shaped by a particular climate and soil. Along the Mediterranean Basin, softer low-growing scrublands carry their own local names: garrigue in France, phrygana in Greece, tomillares in Spain, and batha in Israel. The California coast has its own variant in northern coastal scrub and coastal sage scrub. The Western Cape of South Africa hosts strandveld, and central Chile has coastal matorral. What most of these share is proximity to the sea. Mediterranean scrublands are most common near coastlines and have often adapted to coastal wind and salt air over generations. Interior scrublands follow a different logic, appearing in semi-arid areas with nutrient-poor soils, such as on the matas of Portugal, which sit above Cambrian and Silurian schists. Florida scrub is another inland example of this pattern.

  • Shrubland species carry fire in their biology. Heavy seed production, lignotubers, and fire-induced germination are all adaptations that allow these plants not merely to survive burning but to depend on it. Lignotubers are woody swellings at the base of a stem that store nutrients and allow a plant to resprout rapidly after flames have stripped everything above ground. Fire-induced germination means that certain seeds will not open at all until exposed to heat or smoke, holding their contents in reserve for the moment the canopy clears. Regular natural disturbance by fire or browsing can maintain a shrubland in a stable state indefinitely. Remove that disturbance and the community may shift toward woodland or forest over time. These same adaptations that make the plants resilient create serious risk for people. Shrubland may be unsuitable for human habitation precisely because of the danger of fire. Desert scrublands add further defenses suited to a different stress: small leaves that limit water loss, thorns that deter grazing animals, succulent leaves or stems for storing moisture, and long taproots that reach down to groundwater far below the surface.

  • Not every shrubland is what it appears to be. Some are the climax community for their region, the vegetation that will persist there indefinitely under natural conditions. Others are transitional, a temporary stage that follows a disturbance such as fire and will eventually succeed into something else. Human activity adds a third origin. Logging and overgrazing can degrade forest or woodland until only shrubs remain, and in that case the result is more commonly called shrubbery or bushes rather than shrubland proper. The World Wildlife Fund recognizes multiple shrubland biomes, grouping them into desert scrublands and xeric shrublands, Mediterranean scrublands, and interior scrublands among others. Xeric scrublands appear not only in the world's deserts but also in fast-draining sandy soils within more humid regions, carried there by soil chemistry rather than by rainfall alone. The matas of Portugal illustrate how interior scrublands can be pinned to a place by geology, in that case the ancient Cambrian and Silurian schist underlying the land. Whether a given shrubland is a destination or a way-station depends entirely on what holds it in place, and dwarf-shrub formations such as heathland and moorland, with their acid-loving low-growing species, show that even the smallest and most creeping shrub communities can be stable features of the landscape for a very long time.

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Common questions

What is shrubland and why is it called by so many different names?

Shrubland is a plant community dominated by shrubs and also known as scrubland, scrub, brush, or bush. Regional varieties carry local names such as chaparral in California, matorral in Chile and Spain, maquis in France, fynbos in South Africa, and kwongan in Southwest Australia. The term "shrubland" itself was coined in 1903.

How do botanists define a shrub versus a tree?

A shrub is formally defined in botany and ecology as a much-branched woody plant less than 8 metres high, usually with many stems. Tall shrubs are mostly 2-8 metres high, small shrubs 1-2 metres high, and subshrubs less than 1 metre high.

What fire adaptations do shrubland plants have?

Shrubland species show adaptations including heavy seed production, lignotubers, and fire-induced germination. Lignotubers are woody swellings at the stem base that store nutrients and allow rapid resprouting after fire. Fire-induced germination means some seeds only open when exposed to heat or smoke.

Can shrubland be caused by human activity?

Yes. Shrubland may occur naturally or result from human activity such as logging or overgrazing that degrades former forest or woodland. When it is the product of human disturbance, the terms shrubbery or bushes are more commonly used rather than shrubland.

Why is shrubland sometimes considered unsuitable for human habitation?

Shrubland may be unsuitable for human habitation because of the danger of fire. The fire adaptations of shrubland plants, such as fire-induced germination, mean these ecosystems are structured around burning, which poses significant risks to people living nearby.

What is the Australian vegetation classification system used for shrublands?

Australia uses a descriptive system based on structural characteristics, plant life-form, height, and foliage cover of the tallest stratum. Shrubs of 2-8 metres are classed from closed-shrublands at 70-100 percent foliage cover down to tall open shrubland at under 10 percent cover. A parallel set of categories covers shrubs under 2 metres, ranging from closed-heath to low open shrubland.

All sources

4 references cited across the entry

  1. 1BookEncyclopedia of desertsUniversity of Oklahoma Press — 1999
  2. 2Mediterranean ShrublandsSusan Woodward — Radford University
  3. 3Deserts and Xeric ShrublandsWorld Wildlife Fund