The vascular cambium is the main growth tissue in the stems and roots of plants that exhibit secondary growth, including dicots such as oak trees and gymnosperms such as pine trees. It produces secondary xylem inward toward the pith and secondary phloem outward toward the bark. Generally, more secondary xylem is produced than secondary phloem.
What types of cells make up the vascular cambium?
The vascular cambium consists of two types of cells: fusiform initials, which are tall and axially oriented, and ray initials, which are smaller and round to angular in shape. These two cell types maintain the connection and communication between xylem and phloem.
Which plants do not have a vascular cambium?
Five angiosperm lineages have independently lost the vascular cambium: Nymphaeales, Ceratophyllum, Nelumbo, Podostemaceae, and monocots. Because monocots lack the vascular cambium, plants such as grasses do not undergo secondary growth and cannot develop thick woody trunks.
What hormones regulate the vascular cambium?
The phytohormones involved in vascular cambial activity include auxins, ethylene, gibberellins, cytokinins, abscisic acid, and likely additional hormones yet to be identified. Auxin stimulates mitosis and cell production; gibberellin stimulates cambial cell division and regulates xylem differentiation; cytokinin regulates the rate of cell division.
Why is the vascular cambium important for plant grafting?
For successful grafting, the vascular cambia of the rootstock and scion must be precisely aligned so they can grow together. If the cambial layers do not make contact, the two plant parts cannot merge and the graft fails.
Is the vascular cambium edible and was it ever used as food?
The cambium of most trees is edible. In Scandinavia, it was historically used as a flour to make bark bread.